FRESNO, Calif. — Dane Evans threw four touchdown passes and ran in the game-winning 18-yard touchdown in the second overtime as Tulsa rallied from 31 points down to beat Fresno State 48-41 on Saturday for the program’s largest comeback victory.

Evans was 22 of 32 for 273 yards, Keevan Lucas caught two 40-yard touchdowns, and Josh Atkinson and Justin Hobbs each had scoring catches. D’Angelo Brewer ran for a career-best 252 yards on 46 carries, and James Flanders ran for 101 yards and a touchdown for Tulsa (3-1).

Aaron Peck caught a pass from Chason Virgil on the Bulldogs’ second play of overtime but he fumbled. Then Tulsa’s Redford Jones missed a 42-yard field goal to force a second OT.

Virgil ran for two scores, and threw a 65-yard TD strike to Jamire Jordan to give the Bulldogs a 31-0 lead with 10 minutes left in the second quarter. Virgil threw an incomplete pass on fourth-and-17 to end it.